HOSTA ANALYZED BY FLOW CYTOMETRY

Arranged by Ploidy Level

Diploids (2-2-2)

'Abiqua Drinking Gourd' 2-2-2
'Aki Tenryu' 2-2-2
'Allegan Fog' 2-2-2
'Amanuma' 2-2-2
'Aphrodite' 2-2-2
'Barney Fife' 2-2-2
'Beatrice' 2-2-2
'Bedazzled' 2-2-2
'Bella' 2-2-2
'Belle of the Ball' 2-2-2
'Bill Brincka' 2-2-2
'Blauwichtel' 2-2-2
'Blue Cadet' 2-2-2
'Blue Mouse Ears' 2-2-2
'Blue Splendor' 2-2-2
'Blue Umbrellas' 2-2-2
'Blythe Spirit' 2-2-2
'Botanical Look' 2-2-2
'Breathless' 2-2-2
'Bunter Babette' 2-2-2
'Butter Rim' 2-2-2
'Cadillac' 2-2-2
capitata 2-2-2
'Carolina Sunshine' 2-2-2
'Cascades' 2-2-2
'Cerveza' 2-2-2
'Cheatin Heart'1 2-2-2
'Chelsea Ore' 2-2-2
'Cherry Berry' 2-2-2
'Cherry Tomato' 2-2-2
'Chinese Sunrise' 2-2-2
'Chionea' 2-2-2
'Chishima Iwa' 2-2-2
'Choko Nishiki' 2-2-2
clausa var. normalis 2-2-2
'Climax' 2-2-2
'Collector's Choice' 2-2-2
'Concord' 2-2-2
'Cool as a Cucumber' 2-2-2
'County Park' 2-2-2
'Crispula' 2-2-2
'Crystal Chimes' 2-2-2
'Cynthia' 2-2-2

'Dancing in the Rain' 2-2-2
'Dancing Stars' 2-2-2
'Decorata' 2-2-2
'Dee's Golden Jewel' 2-2-2
'Devon Tor' 2-2-2
'Diamond Tiara' 2-2-2
'Diana Remembered' 2-2-2
'Dorset Blue' 2-2-2
'Dress Blues' 2-2-2
'El Niño' 2-2-2
'Elata' 2-2-2
'Elatior' 2-2-2
'Elegans' 2-2-2
'Embroidery' 2-2-2
'Emerald Necklace' 2-2-2
'Faithful Heart' 2-2-2
'Fall Bouquet' 2-2-2
'Fantabulous' 2-2-2
'Filagree' 2-2-2
"Fire Works"2
'Five O'Clock Shadow' 2-2-2 and 2-3.8-3.8
fluctuans 2-2-2
'Fortunei Albopicta' 2-2-2
'Fourth of July' 2-2-2
'Fragrant Blue' 2-2-2
'Fragrant Bouquet' 2-2-2
'Francee' 2-2-2
'Frances Williams' 2-2-2
'Frosted June' 2-2-2
'Frosted Mouse Ears' 2-2-2
'Fujibotan' 2-2-2
'Gaiety' 2-2-2
'Gaijin' 2-2-2
'Ginko Craig' 2-2-2
'Gold Edger' 2-2-2
'Gold Standard' 2-2-2
'Golden Friendship' 2-2-2
'Golden Memories' 2-2-2
'Golden Tiara' 2-2-2
gracillima 2-2-2
gracillima 'Alba' 2-2-2
'Grand Forks' 2-2-2
'Grand Marquee'3 2-2-2 and 4-2-2
'Grand Slam' 2-2-2
'Grand Tiara'3 4-2-2
'Great Expectations' 2-2-2
'Green Eyes' 2-2-2
'Green Mouse Ears' 2-2-2
'Guacamole' 2-2-2
'Gypsy Rose' 2-2-2
'Hadspen Blue' 2-2-2
'Haku Chu Han' 2-2-2
'Hakuyo' 2-2-2
'Halcyon' 2-2-2
'Happy Dayz' 2-2-2
'Harry van de Laar' 2-2-2
'High Noon' 2-2-2
'Hime Tokudama' 2-2-2
'Hirao Majesty' 2-2-2
'Hirao Supreme' 2-2-2
'Hirao Tetra' 2-2-2
'Holly's Dazzler' 2-2-2
'Honeybells' 2-2-2
'Hudson Bay' 2-2-2
'Hydon Sunset' 2-2-2
'Illicit Affair' 2-2-2
'Inaho' 2-2-2
'Invincible' 2-2-2
'Iron Gate Delight' 2-2-2
'Iron Gate Supreme' 2-2-2
'Iwa Soules' 2-2-2
'Japanboy' 2-2-2
'Japangirl' 2-2-2
jonesii 2-2-2
'June' 2-2-2
'June Fever' 2-2-2
'June Present' 2-2-2
'Kabitan' 2-2-2
'Katsuragawa Beni' 2-2-2
'Key Lime Pie' 2-2-2
'Kifukurin Hyuga' 2-2-2
'Kifukurin Ko Mame' 2-2-2
'Kifukurin Ubatake' 2-2-2
kikutii 2-2-2
kikutii var. caput-avis 2-2-2
kikutii var. leuconata 2-2-2
kikutii var. polyneuron 2-2-2
kikutii var. tosana 2-2-2
kikutii var. yakusimensis 2-2-2
'Kinba' 2-2-2
'Kirishima' 2-2-2
kiyosumiensis 2-2-2
kiyosumiensis 'Busen' 2-2-2
kiyosumiensis 'Maiohgi' 2-2-2
kiyosumiensis 'Seiryu' 2-2-2
'Korean Snow' 2-2-2
'Koryu' 2-2-2
'Lancifolia' 2-2-2
'Lily Pad' 2-2-2
'Liliput' 2-2-2
'Lionheart' 2-2-2
'Little Caesar' 2-2-2
'Little Jay' 2-2-2
'Little Jim' 2-2-2
'Little Wonder' 2-2-2
longipes 2-2-2
longipes 'Amagi Nishiki' 2-2-2
longipes 'Aurea' 2-2-2
longipes f. hypoglauca 2-2-2
longipes 'Gotemba Nishiki' 2-2-2
longipes 'Lancea' 2-2-2
longipes 'Maruba' 2-2-2
longipes 'Ogon' 2-2-2
longipes 'Tenryu Nishiki' 2-2-2
longissima 2-2-2
'Love Pat' 2-2-2
'Maekawa' 2-2-2
'Magic Fire' 2-2-2
'Marbled White' 2-2-2
'Maruba Iwa' 2-2-2
'Mary Marie Ann' 2-2-2
'Masquerade' 2-2-2
'Mata Hari' 2-2-2
'Maui Buttercups' 2-2-2
'Memories of Dorothy' 2-2-2
'Mighty Mouse' 2-2-2
'Mikawa-no-yuki' 2-2-2
minor "Gosan" 2-2-2
montana 2-2-2
montana 'Aureomarginata' 2-2-2
montana f. macrophylla 2-2-2
'Moonstruck' 2-2-2
'Mount Kirishima' 2-2-2
'My Child Insook' 2-2-2
'My Laura' 2-2-2
'Neat Splash' 2-2-2
'Obsession' 2-2-2
'Ogon Tsushima' 2-2-2
'Oh My Heart' 2-2-2
'Old Glory' 2-2-2
'On Stage'4 2-2-2
'Opipara' 2-2-2
'Orange Marmalade' 2-2-2
'Otome-no-ka' 2-2-2
'Outhouse Delight' 2-2-2
'Pandora's Box' 2-2-2
'Paradise Joyce' 2-2-2
'Paul's Glory' 2-2-2
'Piedmont Gold' 2-2-2
plantaginea 2-2-2
'Pocketful of Sunshine' 2-2-2
'Pot of Gold' 2-2-2
'Praying Hands' 2-2-2
pulchella 2-2-2
'Purple Profusion' 2-2-2
pycnophylla 2-2-2
'Rainforest Sunrise' 2-2-2

rectifolia 2-2-2
rectifolia 'Atropurpurea' 2-2-2
rectifolia 'Aurea' 2-2-2
rectifolia 'Sachalin' 2-2-2
rectifolia 'White Tachi' 2-2-2
'Red Dragon' 2-2-2
'Red Neck Heaven' 2-2-2
'Red October' 2-2-2
'Red Tubes' 2-2-2
'Regal Rhubarb' 2-2-2
'Remember Me' 2-2-2
'Rhein' 2-2-2
'Rhino Gold' 2-2-2
rohdeifolia f. viridis 2-2-2
'Rosemore' 2-2-2
'Royal Standard' 2-2-2
'Royal Wedding' 2-2-2
'Rumpelstilzchen' 2-2-2
rupifraga 2-2-2
'Sagae' 2-2-2
'Saikoku' 2-2-2
'Saint Elmo's Fire' 2-2-2
'Saishu Jima' 2-2-2
'Setsurei' 2-2-2
'Shelleys' 2-2-2
'Sherborne Swift' 2-2-2
shikokiana 2-2-2
'Shining Tot' 2-2-2
sieboldiana 2-2-2
sieboldii 2-2-2
sieboldii 'Alba' 2-2-2
'Silver Eagle' 2-2-2
'Silver Threads & Golden Needles' 2-2-2
'Sir Richard' 2-2-2
'Sitting Pretty' 2-2-2
'Sleeping Beauty' 2-2-2
'Slick Willie' 2-2-2
'Snow Cap' 2-2-2
'Snow Flakes' 2-2-2
'Snow Mouse' 2-2-2
'Snow White' 2-2-2
'Sparkling Burgundy' 2-2-2
'Sparky' 2-2-2
'Spilt Milk' 2-2-2
'Stained Glass' 2-2-2
'Stand by Me' 2-2-2
'Stargate' 2-2-2
'Steady On' 2-2-2
'Stiletto' 2-2-2
'Straka Gold' 2-2-2
'Striptease' 2-2-2
'Swan Song' 2-2-2
'Sweet Susan' 2-2-2
takahashii "Gosan" 2-2-2
'Tardiflora' 2-2-2
'Tardiflora Hybrida' 2-2-2
tardiva "Gosan" 2-2-2
'Tea Time' 2-2-2
'Tetra' 2-2-2
'Theo's Blue' 2-2-2
tibae 2-2-2
'Tiny Tears' 2-2-2
'Tokudama' 2-2-2
'Tokudama Aureonebulosa' 2-2-2
'Tokudama Flavocircinalis' 2-2-2
'Torchlight' 2-2-2
'Treasure Island' 2-2-2
'Tsugaru Komachi' 2-2-2
tsushimensis 2-2-2
'Twilight' 2-2-2
'Twilight Zone' 2-2-2
'Ultraviolet Light' 2-2-2
'Undulata Albomarginata' 2-2-2
'Undulata Erromena' 2-2-2
'Undulata Univittata' 2-2-2
'Urajiro Hachijo' 2-2-2
'Urajiro Hachyo' 2-2-2
'Uzu-no-mai' 2-2-2
venusta 2-2-2
'Vera Verde' 2-2-2
'Victory' 2-2-2
'Viridipes' 2-2-2
'Warwick Comet' 2-2-2
'Waukon Thin Ice' 2-2-2
'Wheee!' 2-2-2
'White Bikini' 2-2-2
'White Tachi' 2-2-2
'White Wall Tire' 2-2-2
'William Lachman' 2-2-2
'Windsor Blue' 2-2-2
'Yakushima Mizu' 2-2-2
'Yankee Blue' 2-2-2
'Yea' 2-2-2
yingeri 2-2-2
'Yoshinogawa' 2-2-2
'Yu Lei' 2-2-2

Triploids (3-3-3)

'Amine Tachi' 3-3-3
'Blazing Saddles' 3-3-3
'Blizzard' 3-3-3
'Bottom Line' 3-3-3
clausa var. clausa 3-3-3
'Corona' 3-3-3
'David A. Haskell' 3-3-3
'Domain de Courson' 3-3-3
'Fran Godfrey' 3-3-3
'Guardian Angel' 3-3-3
'Gunther's Prize' 3-3-3
'Holly's Honey' 3-3-3
'Lady Isobel Barnett' 3-3-3
'Lakeside Coal Miner' 3-3-3
'Little Blue'5 3-3-3
'Parhelion' 3-3-3
'PeeDee Elfin Bells' 3-3-3
'Prairie Sum Shine' 3-3-3
'Shiny Substance' 3-3-3
'Sum and Substance' 3-3-3
'Sum and Subtle' 3-3-3
'Sum Cup-o-Joe' 3-3-3 and 2.8-2.8-2.8
'Sum It Up' 3-3-3
'Tiffney's Godzilla' 3-3-3
'Tucker Valentine' 3-3-3
'Venucosa' 3-3-3
'Winter Snow' 3-3-3
'Zebson' 3-3-3

Tetraploids (4-4-4)

'Amber Tiara' 4-4-4
'Bulletproof' 4-4-4
'Cathedral Windows' 4-4-4
'Christmas Candy' 4-4-4
'Dance with Me' 4-4-4
'Doubled Up' 4-4-4
'Dust Devil' 4-4-4
'Eternal Flame' 4-4-4
'Final Victory' 4-4-4
'Fire and Ice' 4-4-4
'Forbidden Fruit' 4-4-4
'Frosted Dimples' 4-4-4
'Golden Needles' 4-4-4
'Grand Tiara'3 4-2-2 and 4-4-4
'Hands Up' 4-4-4
'Independence' 4-4-4
'Jolly Green Dwarf' 4-4-4
'Justine' 4-4-4
'Kiwi Black Magic' 4-4-4
'Lakeside Black Satin' 4-4-4
'Lettuce Salad' 4-4-4
'Loyalist' 4-4-4
'Minuteman' 4-4-4
'Mojito' 4-4-4
'Morning Light' 4-4-4
'My Friend Bertha' 4-4-4
'Neutron Star'6 4-4-4
'Orange Star' 4-4-4
'Otome' 4-4-4
'Pathfinder' 4-4-4
'Patriot'3 4-2-2 and 4-4-4
'Pitch Black' 4-4-4
'Prince of Darkness' 4-4-4
'Radiant Edger' 4-4-4
'Revolution' 4-4-4
'Rhino Blue' 4-4-4
'Second Wind' 4-4-4
'Second Wind' 4-4-4
'Secret Ambition' 4-4-4
'Shade Master' 4-4-4
'Summer Breeze' 4-4-4
'Summer Lovin' 4-4-4
'Touch of Class'3 4-2-2 and 4-4-4
'Touch of Frost' 4-4-4
'Trade Wind' 4-4-4
'Twilight' 4-4-4
ventricosa 4-4-4
ventricosa 'Aureomaculata' 4-4-4
ventricosa 'Aureomarginata' 4-4-4
ventricosa 'Golden Spot' 4-4-4
ventricosa 'Schwarzer Ritter' 4-4-4
ventricosa 'Spring Gold' 4-4-4
'Volcano Island' 4-4-4
'Warwick Essence' 4-4-4
'Whirlwind' 4-4-4
'Woop Woop' 4-4-4

Ploidy Chimeras

'Atlantis' 4-2-2
'Avocado' 4-2-2
'Blue Ivory' 4-2-2
'Captain Kirk' 4-2-2
'Dream Weaver' 4-2-2
'Enterprise' 4-2-2
'First Light' 4-4-6
'Goodness Gracious' 4-2-2
'Grand Marquee'3 2-2-2 and 4-2-2
'Grand Tiara'3 4-2-2 and 4-4-4
'Great Escape' 4-2-2
'Heaven Scent' 4-2-2
'Iris Frazier' 3-2-2
'Ivory Coast' 4-2-2
'Last Dance' 3-3-4
'Liberty' 4-2-2
'Linda Sue' 4-2-2
'Little Treasure' 4-2-2
'Majesty' 4-2-2
'Middle Ridge' 4-2-2
'Mourning Dove' 4-2-2
'Night before Christmas' 4-2-2
'Olympic Sunrise' 4-2-2
'Patriot'3 4-2-2 and 4-4-4
'Rhino Hide' 4-2-2
'Royal Super' 2-2-4
'St. Paul' 4-2-2
'Summer Music' 4-2-2
'Sweet Innocence' 4-2-2
'Touch of Class'3 4-2-2 and 4-4-4
'Vulcan' 4-2-2

Aneuploids

'Academy Ametrine' Aneuploid
'After Midnight' 3.8-2-2
'Bubba'7 Aneuploid
'Buster' 3.4-3.4-3.4 and 3.5-3.5-3.5
'Eagle's Nest' 3-3-4.3
'Final Summation' 2.9-2.9-2.9
'Fireworks' 3.2-4-4
'Five O'Clock Shadow' 2-2-2 and 2-3.8-3.8
'Five O'Clock Somewhere' 3.8-2-2
'Follow Your Dream' 2.9-2.9-2.9
'Fragrant Gold' Aneuploid
'Friar Tuck' 3.5-3-3
'Grand Prize' 3.6-2-2
'Jaz' 2.4-2.4-2.4
'Jim Cooper' Aneuploid
'Joyce Trott' Aneuploid
'Leather Sheen' Aneuploid
'Lodestar' 2.7-2.7-2.7
'Lovely Loretta' Aneuploid
'M. Owen Papuga' Aneuploid
'Mango Tango' 4-2.8-4
'Metallic Sheen' Aneuploid
'Nancy Gill' 2.7-2.7-2.7 and 2.8-2.8-2.8
'New Wave' Aneuploid
'Prestige and Promise' 3.8-3.8-2
'Rainbow's End' 2.5-2-2 and 2-2-2.5
'Raleigh Remembrance' 2.5-2.5-2.5
'Rosedale Genevieve' Aneuploid
'Small Sum' Aneuploid
'Something Good' 3-2.7-2.7
'Sparkler' 3.2-3.2-3.2
'Stitch in Time' 4-2.5-3.6
'Sum Cup-o-Joe' 3-3-3 and 2.8-2.8-2.8
'Sum of All' 3-2.7-2.5
'Titanic'8 3-2.8-2.8
'Tossed Salad' 2.3-2.3-2.3
'True North' 2.9-2.9-2.9
'Vim and Vigor' 2.8-2.8-2.8
'Zero Sum' 2.9-2.9-2.9

1According to Steven C. Chamberlain, "Ploidy and Pollen Viability in Hosta Cultivars: Zonneveld's Data Sets," The Hosta Journal 42, no. 1 (2011): 71-78, H. 'Cheatin Heart' is presented as a ploidy chimera (2-2-4) and a diploid (2-2-2). Upon re-examination of the flow cytometry data, it was determined that 'Cheatin Heart' is diploid (2-2-2).

2Ben J. M. Zonneveld and Frank Van Iren, "Flow cytometric analysis of DNA content in Hosta reveals ploidy chimeras", Euphytica, 111, no. 2 (2000): 105-110 lists H. "Fire Works" as diploid. This hosta is not H. 'Fireworks' PP16,062 (Walters Gardens- 2001). Zonneveld and Van Iren, "H. 'Fireworks' and 'Sparkler': Unusual Aneuploid Sports from 'Loyalist'," The Hosta Journal 46, no. 2(2015): 72-76 states that H. 'Fireworks' (Walters Gardens- 2001) is aneuploid, 3.2-4-4.

3According to Zonneveld and Frank Van Iren, "Flow cytometric analysis of DNA content," H. 'Patriot' is tetraploid. However, B. J. M. Zonneveld, "Nuclear DNA content of ploidy chimeras of Hosta Tratt. (Hostacae) demonstrate three apical layers in all organs, but not in the adventitious root,"Plant Systematics and Evolution 269, no. 1 (2007): 29-38 states that H. 'Patriot' is partially tetraploid (4-2-2). Eventually, it was discovered that both partially tetraploid as well as fully tetraploid forms are in the trade. According to B. J. M. Zonneveld and Warren I. Pollock, "Flow cytometric analysis of somaclonal variation in lineages of Hosta sports detects polyploidy and aneuploidy chimeras," Plant Biology 14, no. 6 (2012): 972-979, "H. 'Patriot' exists as both partial tetraploid (4-2-2) and fully tetraploid (4-4-4) plants in the trade (Zonneveld 1999). Several other plants with different ploidy are in commerce under the same name. Examples are H. 'Grand Tiara', H. 'Grand Marquee' and H. 'Touch of Class', with both 4-2-2 and 4-4-4 in the trade. Seemingly, over the years, tissue culture propagation of 4-2-2 plants has given rise to 4-4-4 plants. As 4-2-2 plants have leaves comparable in thickness to 4-4-4 plants, this change might go unnoticed." Also see, Ben J. M. Zonneveld, "The DNA Content of H. 'Francee', 'Minuteman', and Two Different Varieties Called 'Patriot'," The Hosta Journal 30, no. 2 (1999): 28, 29.

4According to Warren I. Pollock, "Reflections: 2008," The Hosta Journal 40, no. 1 (2009): 72-76, H. 'On Stage' is incorrectly presented as tetraploid. In Zonneveld and Van Iren, "Flow cytometric analysis of DNA content," H. 'On Stage' is confirmed as diploid.

5According to Zonneveld and Frank Van Iren, "Flow cytometric analysis of DNA content," H. 'Little Blue' is incorrectly identified as a tetraploid (page 3) and an aneuploid (page 5). Zonneveld has since determined H. 'Little Blue' to be a triploid.

6Hosta 'Neutron Star' was originally submitted as H. 'Otome' for flow cytometry measurement. See Steven C. Chamberlain, Warren I. Pollock, and Ben J. M. Zonneveld, "Ploidy and Pollen Viability in Hosta Cultivars: Zonneveld's Data Sets", The Hosta Journal 42, no. 1 (2011): 71-78. The hosta was determined to be 4-4-4 and the name was later changed to 'Neutron Star'.

7According to Warren I. Pollock, "This and That: 2011," The Online Hosta Journal 42 Online, H. 'Bubba' is aneuploid, not diploid as previously stated by Steven C. Chamberlain, Warren I. Pollock, and Ben J. M. Zonneveld, "Ploidy and Pollen Viability in Hosta Cultivars: Zonneveld's Data Sets," The Hosta Journal 42, no. 1 (2011): 71-78.

8Steven C. Chamberlain, Warren I. Pollock, and Ben J. M. Zonneveld, "Ploidy and Pollen Viabiliy in Hosta Cultivars: Zonneveld's Data Sets," The Hosta Journal 42, no. 1 (2011): 71-78 incorrectly presents H. 'Titanic' as triploid. However, Zonneveld and Pollock, "Flow cytometric analysis of somaclonal variation," confirms H. 'Titanic' is aneuploid, 3-2.8-2.8.

Sources:

Chamberlain, Steven C. "Zonneveld's Study of Cell Layers and Leaf Variegation in Hosta Cultivars: A Summary."
    The Hosta Journal, Volume 39, Number 2 (2008): 66-69.

Chamberlain, Steven C., Warren I. Pollock, and Ben J. M. Zonneveld. "Ploidy and Pollen Viability in Hosta Cultivars: Zonneveld's Data Sets."
The Hosta Journal, Volume 42, Number 1 (2011): 71-78.

Mortko, Rob. "Hot Hostas, Chemically Induced Tets: A Sure Fire Method for Success?"
   The Online Hosta Journal, Volume 41 Online (2010).

Pollock, Warren I. Email to Donald Rawson regarding H. 'Majesty' and H. 'Victory'. February 21, 2015.

Pollock, Warren I. "Reflections: 2008." The Hosta Journal, Volume 40, Number 1 (2009): 72-76.

Pollock, Warren I. "This and That: 2011." The Online Hosta Journal, Volume 42 Online (2011).

Pollock, Warren I. "This and That: 2015; New Hosta Ploidy (L1-L2-L3) Measurements."
    The Online Hosta Journal, Volume 46 Online (2015).

Pollock, Warren I. "What's in a Hosta Name? Part LIII." The Hosta Journal, Volume 42, Number 2 (2011): 67-74.

Pollock, Warren I. "What's in a Hosta Name? Part LVII." The Hosta Journal, Volume 44, Number 2 (2013):8-18.

Pollock, Warren I and Ben J. M. Zonneveld. "More Information on Bob Solberg's 'Hybridizing thoughts 2015'."
   The Online Hosta Journal, Volume 46 Online (2015).

Solberg, Bob. "Do You Know a Tetraploid Hosta When You See One?" The Hosta Journal, Volume 42, Number 2 (2011): 33-36.

Solberg, Bob. Green Hill Farm. <http://www.hostahosta.com>.

Walek, Kevin. "Neutron Star". The Hosta Registry. 2011. <http://www.hostaregistrar.org>.

Zonneveld, Ben J. M. "Nuclear DNA content of ploidy chimeras of Hosta Tratt. (Hostaceae) demonstrate three apical layers in all organs, but not in the
    adventitious root." Plant Systematics and Evolution, Volume 269, Issue 1 (2007): 29-38.

Zonneveld, Ben J. M. "The DNA Content of H. 'Francee', 'Minuteman', and Two Different Varieties Called 'Patriot'."
    The Hosta Journal, Volume 30, Number 2 (1999): 28, 29.

Zonneveld, Ben J. M. and Frank Van Iren. "Flow cytometric analysis of DNA content in Hosta reveals ploidy chimeras."
   Euphytica, Volume 111, Issue 2 (2000): 105-110.

Zonneveld, Ben J. M. and Frank Van Iren. "Genome Size and Pollen Viability as Taxonomic Criteria: Application to the Genus Hosta."
    Plant Biology, Issue 3 (2001): 176-185.

Zonneveld, Ben J. M., Warren I. Pollock, Rob Mortko, and Steven C. Chamberlain. "H. 'Stitch in Time': A Multiple Mystery."
    The Hosta Journal, Volume 40, Number 1 (2009): 62-71.

Zonneveld, Ben J. M. and Warren I. Pollock. "Flow cytometric analysis of somaclonal variation in lineages of Hosta sports detects polyploidy
    and aneuploidy chimeras." Plant Biology, Volume 14, Issue 6 (2012): 972-979.

Zonneveld, Ben J. M. and Warren I. Pollock. "H. 'Fireworks' and 'Sparkler': Unusual Aneuploid Sports from 'Loyalist'."
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